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The field may contain magenta
text presenting data from a type study
and/or revision of other original material cited in
the protolog of the present taxon.
Macroscopic descriptions in magenta are a
combination of data from the protolog and
additional observations made on the exiccata during
revision of the cited original
The same field may also contain black text, which is
data from a revision of the present
taxon (including non-type material and/or material
not cited in the protolog).
Paragraphs of black text will be labeled if further
this text is appropriate.
Olive text indicates a specimen
that has not been
thoroughly examined (for example, for microscopic
details) and marks other places in the text
where data is missing or uncertain.
The following material is based molecular research
of Dr. Linas Kudzma and other original research of
R. E. Tulloss.
62 mm wide, dark brown over disc, elsewhere paler brown,
unchanging when cut or bruised,
broadly planoconvex, with small central umbo, tacky,
margin nonappendiculate, tuberculate-strate
(0.40R); context pale sordid white, staining
not observed,4.5 mm thick above stipe, thinning evenly
toward margin for 65 - 75% of gill length, then
membranous to margin; universal veil absent.
free, with very short decurrent tooth (10× lens),
subcroweded to crowded, pale cream in mass, white in
side view, with bruising not oberved, 6 mm broad;
lamellulae truncate, unevenly distributed,
plentiful, of diverse lengths.
108 × 8 mm, white to off-white,becoming pale
brownish from handling, narrowing upward, just barely
flaring at apex, smooth for upper two-thirds, with
finely fibrillosed raised squamules in lower third above
volval limb; context hollow, off-white to very
pale yellowish white, with central cylinder 3 mm wide,
invertebrate damage not present;
exannulate; universal veil saccate,
persistent, soft, membranous, 14.5 mm wide, 41 mm
from very base of stipe to highest point of limb,
less than 1 mm thick, subabruptly flaring away from
stipe at about mid-height, below this point,
appressed to stipe, dominately white on both
surfaces, with pale orangish brown patches on
expanded upper portion of limb, appressed to stipe
one half height; limbus internus arising from
about point of separation of stipe and universal
veil, paper thin except for short basal portion
having triangular cross-section.
Odorless and tastless.
Spot test for laccase (syringaldazine) - rapidly
positive in spot below cental cavity of stipe,
spreading along line dividing tissues of stipe base
from limb of universal veil; otherwise, entirely
negative. Spot test for tyrosinase (paracresol) - positive
throughout basidiome except for central portion of
pileus context, spots on lamellae, and context of
universal veil at stipe base. Test voucher: