This species is dominantly white or whitish with some tint of pink or pallid orange at first. The spores are subglobose to broadly ellipsoid.
The cap of Amanita homolae is 52 - 78 mm wide, cream or very pale pinkish- or orangish-white, becoming sordid yellowish cream in age, broadly campanulate with a deflexed margin, becoming planoconvex, tacky, dull, with a striate margin (35 - 60% of the radius), nonappendiculate. There are remains of a volva on the cap. The flesh is white, unchanging when cut or bruised, 4.5 - 6.5 mm thick above the stem, thinning evenly for one half to two thirds of radius, then membranous to the margin.
The gills are free to narrowly adnate, subcrowded to crowded, pale pinkish- or orangish-cream in mass, drying a lovely pale orange, 6.5 or more mm broad, with a decurrent tooth or a line on the top of the stem. The short gills are truncate, of varying length, and unevenly distributed.
The stem is 140 - 220 × 12-13.5 mm, entirely whitish or cream with an orange tint in the upper half and dingy gray above the saccate volva, browning on the edges of wounds and from handling. The stem is ringless. The saccate volva is white, occasionally with some brown stains on the exterior surface, membranous, leathery, free from the stem for considerably more than half the distance from the stem base to the highest point on the limb, about 1 mm thick at the midpoint between the highest point of the limb and the point of attachment to the stem, with the highest point on a limb 49 - 51 mm from the base of the stem. The limbus internus is present as a very small ridge just above point of attachment to the stem. The flesh is white to off-white and sometimes somewhat water logged, unchanging when cut or bruised, hollow or partially stuffed.
An odor for this species has not been reported
The spores measure 9.2 - 11.0 × (8.2-) 8.5 - 9.5 µm and are globose to subglobose to broadly ellipsoid and inamyloid. There are no clamps at the bases of basidia.
This species was previously referred to as Amanita sp. N26 in keys and checklists.—R. E. Tulloss and L. Possiel
Tulloss nom. prov.
genitive of a Latinized name; hence "Homola's" or "of Homola"
Due to delays in data processing at GenBank, some accession numbers may lead to unreleased (pending) pages.
These pages will eventually be made live, so try again later.
17.viii.2001 Terry Stoleson s.n. [Tulloss 8-17-01-E] (RET 361-4)
L. Nagy, Agaricales Diversification, Szeged Univ.
Olive text indicates a specimen that has not been
thoroughly examined (for example, for microscopic details) and marks other places in the text
where data is missing or uncertain.
The following text is based on original research of R. E. Tulloss.
52 - 78 mm wide, cream or very pale pinkish- or
orangish-white, or off-white becoming sordid yellowish
cream or pallid
grayish brown in age, broadly campanulate with
deflexed margin, becoming planoconvex, tacky,
dull; context white, whiter than lamellae,
unchanging when cut or bruised, 4.5 - 6.5 mm thick at
stipe, thinning evenly for one half to two thirds of
radius, then membranous to margin; margin
striate (0.35R - 0.6R), nonappendiculate; universal
free to narrowly adnate with a decurrent tooth (lens) or a line on the stipe apex (lens), subcrowded to crowded, pale pinkish- or orangish-cream in mass, ?? in side view, drying a lovely pale orange (as does, on occasion, the underside and striae of the pileus), 6.5 - ?? mm broad; lamellulae truncate, of varying length, unevenly distributed.
140 - 220 × 12-13.5 mm, entirely whitish or cream with orange tint in upper half and dingy gray above the saccate volva, browning on edge of wounds and from handling; context white to off-white and somewhat water logged, unchanging when cut or bruised, larva tunnels concolorous, hollow or partially stuffed with white cottony material, with central cylinder 3.5 - 6.5 mm wide; exannulate; universal veil as saccate volva, white, occasionally with some brown stains on exterior surface, membranous, leathery, free from stipe for considerably more than half distance from stipe base to highest point on limb, about 1 mm thick at midpoint between highest point of limb and point of attachment to stipe, with highest point on limb 49 - 51 mm from base of stipe; limbus internus as a very small ridge just above point of attachment to stipe.
Amanita homolae is a very distinctive species with
its cream-colored cap tinted with pink or orange and a
stipe often decorated with pale orange
Genetic studies indicate that the number of white- to
cream-capped species with a relatively persistent,
saccate universal veil in Amanita section
Vaginatae in northeastern North America are far
more numerous than has been previously suspected.
Due to recent efforts of members of the Quebec and
Montreal mycological societies, the laboratory of Dr.
Jean-Marc Moncalvo, and the authors of this page, the
following white or pallid taxa (in additions to the
present species) have been genetically segregated:
At present, an additional 4?? probable taxa (including A. sp-NFL02) have been identified based on RET's herbarium collections which had been tentatively identified under one of the names listed above.
Genetic work is being scheduled to compare this taxon with
A. sp-NFL02 and A. littenii.
—R. E. Tulloss and L. V. Kudzma
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