The following text may make multiple use of each data field.
The field may contain magenta text presenting data from a type study
and/or revision of other original material cited in the protolog of the present taxon.
Macroscopic descriptions in magenta are a combination of data from the protolog and
additional observations made on the exiccata during revision of the cited original
The same field may also contain black text, which is data from a revision of the present
taxon (including non-type material and/or material not cited in the protolog).
Paragraphs of black text will be labeled if further subdivision of
this text is appropriate.
Olive text indicates a specimen that has not been
thoroughly examined (for example, for microscopic details) and marks other places in the text
where data is missing or uncertain.
The following material is based on the 2006
publication of Yves Lamoureux, notes and
photographs of the diverse collectors and RET, on
molecular research by Dr. Linas V. Kudzma, and
other original research by Dr. Jacques Landry and
80 - 120 mm wide, olivaceous yellow to brownish
yellow, sometimes darker (to umbrinous) over the
umbo and in broad zone around umbo (but not
overlapping marginal striations), at first ovoid,
then rounded-conic, finally planar with distinct
umbo; context white; margin
nonappendiculate, striate (0.2 - 0.25R);
universal veil absent.
RET:47 mm wide, brown over disc, otherwise tan to pinkish
tan, distinctly to
indistinctly virgate, unchanging when bruised or cut,
irregular, tacky, subshiny, not becoming viscid when
moistened; contextnearly white, 4.5 mm thick
above stipe, thinning evenly for 80% of gill length,
then membranous to margin;
margin nonappendiculate, striate (0.15 -
0.30R); universal veil absent.
RET: free to receding, without decurrent tooth on stipe,
crowded, off-white to pale cream
in mass, off-white in side view, unchanging when cut
or bruised, 4 mm broad, broadest at 65-70% of
lamella length from stipe; lamellulae
subtruncate, of diverse lengths, plentiful.
Lamoureux (2006): 200 - 300 × 10 - 20 mm, with pallid base color, narrowing upward, decorated with fibrils (concolorous with pileus) organized in zebroid pattern; context white; exannulate; universal veil as saccate volva, white on exterior, membranous, persistent, 49± × 21± mm, with zigzag upper edge of free limb, with limbus internus not described.
RET: ?? × 8 mm, white,
narrowing upward, flaring at apex, with some fine
raised surface fibrils darkening (to gray or nearly
in upper half, finely striatulate; context
nearly white, unchanging when cut or bruised,
partially stuffed with white cottony material
unevenly distributed, with larval tunnels not
observed, with central cylinder 2.5 mm wide;
exannulate; universal veil membranous,
Odor indistinct. Taste not
RET: Odor indistinct. Taste not
RET: 53- - 59+ × 12.4 - 19.3 μm,
with broadest basidia 2-sterigmate and in mature
regions of hymenium 4-sterigmate and in
immature regions of hymenium commonly 2-sterigmate;
??; clamps not observed.
RET: [36/1/1] (9.2-) 9.6 - 11.0 (-11.3) × (8.2-)
8.5 - 10.0 (-10.4) μm, (L = 10.2 μm; W =
9.1; Q = (1.05-) 1.07 - 1.22 (-1.27); Q =
1.13), hyaline, colorless, smooth, thin-walled,
inamyloid, subglobose to broadly ellipsoid, adaxially
flattened; apiculus sublateral, cyindric;
contents multiguttulate with or without small
granules; color in deposit not recorded.
Solitary. In forests of Fagus or
Quercus. July to September, rare.
Solitary. Canada—Québec: Under
USA—At ca. 72 m elev. Connecticut:
In black loam of mixed deciduous-coniferous
forest. Illinois: In
New Jersey: in wet loam of mixed deciduous forest
with Acer, Betula, Carpinus
caroliniana, Carya ovata, Fagus
grandifolia, Liquidambar styracifluae,
Pennsylvania: in mixed hardwood forest.
12.vii.2008 Renée Lebeuf 93 (RET 447-6, nrITS
& nrLSU seq'd.).
La-Vallée-de-Richelieu - Mont-Saint-Hilaire,
9.vii.1993 G. K. & Y. Lamoureux 1956 (CFFM;
RET 521-9, nrITS seq'd.).
CONNECTICUT—Middlesex Co. - East
Haddam, Devil's Hopyard St. Pk. [41.4756° N/
72.3403° W, 72 m], 1.viii.2015
Mike Rapp s.n. [Tulloss 8-1-15-D] (RET 703-8,
nrITS & nrLSU seq'd.).
ILLINOIS—Monroe Co. - Waterloo, 11.vi.2014
Patrick Harvey 1 (RET 621-10, nrITS seq'd.).
NEW JERSEY—Warren Co. - Stephens St.
Pk., ca. lower picnic area, 4.viii.1996 R. E. Tulloss
8-4-96-D (RET 229-3, nrITS seq'd.)
PENNSYLVANIA—Luzerne Co. - Ricketts Glen
St. Pk. [41.3036° N/ 76.2740° W, 400-600 m],
7.viii.2012 David Wasilewski
(RET 524-10, nrITS & nrLSU seq'd.), 12.vii.2014
D. Wasilewski s.n. [mushroomobserver
(RET 642-9, nrITS & nrLSU seq'd.).
Lamoureux (2006) remarks
that he had only encountered material of this taxon
at two locations in Québec in addition to the one
cited above: Longueuil (Montérégie) and Eastman
(Estrie). He also notes that because the stipe
is deeply inserted in the substrate considerable care
is require to collect a basidiome unbroken.
Deep insertion of the stipe in the substrate occurs
in a number of species of section Vaginatae;
however, this shared character does not imply
a close relationship between the taxa in
question. For example, the present species
seems rather distant from A.
Geoffrey Kibby is cited as having seen this species
in New Jersey.
There are four available nrLSU sequences
numbers provided under "material examined," above)
attributable to the proposed Amanita sp-N65.
All three have the following form: Final
characters (up to 10) of the 3' end of ITS2 are
placed at the
5' end of the sequence. In this region there is
no difference between the sequences. In their
alignment the 5'
region of the LSU sequence begins at character
eleven (11). The 3' terminal character position of
of the three sequences are as follows:
RET 703-8: 1484
RET 642-9: 1495
RET 524-10: 1494
RET 447-6: 1462
To take geographical location into account,
RET 703-8 is the sole Connecticut specimen; the sole
Canadian collection is RET 447-6; and
the other two are from Luzerne County,
The evidence for a SNP (single nucleotide
polymorphism) is provided when
there is variation at a single position between
the two or more sequences in an alignment.
The character positions at which variation occurs
among the four sequences are listed below showing
the variation occurring in each case.
The letter codes indicating ambiguities in the
original sequence data are IUPAC codes for ambiguities
Each spore data set is intended to comprise a set of measurements from a single specimen made by a single observer;
and explanations prepared for this site talk about specimen-observer pairs associated with each data set.
Combining more data into a single data set is non-optimal because it obscures observer differences
(which may be valuable for instructional purposes, for example) and may obscure instances in which
a single collection inadvertently contains a mixture of taxa.