22.ix.2002 P.-A. Moreau s.n. (holotype, in herb. P. Neville 02.09.19.11a => in herb. S. Poumarat
Fac. Pharm., Univ. Lille
in herb. P. Neville 02.09.19.11a => in herb.
S. Poumarat => G; isotype, in herb. P. Neville
02.09.19.11b => in herb. S. Poumarat => G
The following text may make multiple use of each data field.
The field may contain magenta text presenting data from a type study
and/or revision of other original material cited in the protolog of the present taxon.
Macroscopic descriptions in magenta are a combination of data from the protolog and
additional observations made on the exiccata during revision of the cited original
The same field may also contain black text, which is data from a revision of the present
taxon (including non-type material and/or material not cited in the protolog).
Paragraphs of black text will be labeled if further subdivision of
this text is appropriate.
Olive text indicates a specimen that has not been
thoroughly examined (for example, for microscopic details) and marks other places in the text
where data is missing or uncertain.
The following material is derived from the protolog
of the present species, Hanss & Moreau
a field trip in Norway organized by Dr. Gro Gulden,
molecular studies by B. Dima and S. D. Russell, and
other original research by R. E. Tulloss.
58 mm wide, brown with pigment expanding to the formerly
pallid margin after several hours, rounded concave, very
viscid when wet, becoming subshiny upon return to lab;
context 6 mm thick, thinning evenly for 90% of
lamella length, then membranous to cap margin;
margin striate (0.25 - 0.3R), incurved at
first, not appendiculate; universal veil
off-white to orangish white, as thick plaques of
membranous material, smooth with edges curling upward
during drying in wrapper, easily removed, becoming
grayish or (at least) a bit sordid in age.
free or very narrowly adnate, pale brownish gray in
mass due to brown marginal pigmentation, crowded, with
color in side view and staining or
bruising reaction not recorded, like ??
72 × 7.5 mm, with pallid ground covered with
small brown flecks or fibrils except for last 7-8 mm
above join with universal veil giving stem appearance
of having "striangulate" basal zone bordered above
with a thin wavy line of densely place brown fibrils,
slightly browner where handled, narrowing upward,
flaring slowly over ca.
5 mm at apex, longitudinally striatulate;
pale brownish with pale orangish tint in base,
with moderately dense longitudinally oriented white
fibrillose material, with larval tunnels not observed;
exannulate; universal veil as smooth
saccate membranous volva, 33 × 23 mm, about 1 mm
at mid-point between top of volval limb and
attachment of limb to stipe, flaring with multiple
lobes, with exterior surface
whitish, with interior surface palely concolorous
pileus; limbus internus rather high on stem
firmly attached to stem when observed, with
impression of marginal striations visible in strongly
joining outer surface of limbus internus with inner
surface of primary volval limb.
Odor indistinct. Taste not recorded.
from protolog: .
lamella edge tissue
Note: not measured and reported correctly in
protolog—values of Q commonly < 1.0.
Cannot be compared with other taxa.
from protolog: Under
Betula pendula in lawn or under mixed
Betula and Salix.
[Note: Paratype collections were
made in environments in which other genera of
trees were present. This data field will be
RET: Norway: In mineral soil and loam with
Betula near meadow or in wet Betula
forest with herbaceous understory.
Sweden: In poor, sandy ground of grassy path near
Salix myrsinifolia, S. phylicifolia,
S. caprea, S. borealis & Betula
HAUTE-LOIRE—la Chaise-Dieu, 22.ix.2002
P.-A. Moreau s.n. (holotype, in herb. P.
Neville 02.09.19.11a => in herb. S. Poumarat;
isotype, in herb. P. Neville 02.09.19.11b =>
in herb. S. Poumarat => G).
[Note: Paratype collections
remain to be added.—ed.]
21.viii.1999 P. Marstad, H. Myrhe, T. Torjesen s.n.
[Tulloss 8-21-99-A] (O 63282; RET 308-7, nrITS & nrLSU seq'd.),
P. Marstad, H. Myhre s.n. [Tulloss 8-21-99-D]
(O 63283; RET 278-4, nrITS-LSU seq'd.).
Stor-Ängesbäcken, 23.viii.2009 Irene Andersson
(RET 454-9, nrITS-LSU seq'd.).
This taxon was previously called "A. sp-NOR01"
on this site.
—R. E. Tulloss
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can be found here.
2. Amanita betulae, la Chaise Dieu, Haute Loire, France. (type)
Irene Andersson - (1) Stor-Ängesbäcken, Heligfjäll,
Vilhelmina, Lapland, Sweden.
(RET 454-9) [Note: original
unedited images can be found
Dr. Pierre-Arthur Moreau - (2) la Chaise Dieu, Haute
Loire, France. (type) [Note:
reproduced by permission of Dr. Moreau.—ed.]
Spore data for collections provisionally identified as: Amanita betulae Neville & Poumarat
Each spore data set is intended to comprise a set of measurements from a single specimen made by a single observer;
and explanations prepared for this site talk about specimen-observer pairs associated with each data set.
Combining more data into a single data set is non-optimal because it obscures observer differences
(which may be valuable for instructional purposes, for example) and may obscure instances in which
a single collection inadvertently contains a mixture of taxa.